Lycopodiaceae

HUPERZIA Bernh.

  1. Huperzia australiana (Herter) Holub, Folia Geobot. Phytotax. 20: 70 (1985); Sun et al., Plants 13 (2024); POWO (2026). = Borneo, Sulawesi, and SE Australia. = KALIMANTAN, SULAWESI. = Lycopodium australianum Herter in Beibl. Bot. Jahrb. Syst. 98: 34, 48 (1909).
  2. Huperzia beccarii (Alderw.) Holub, Folia Geobot. Phytotax. 20: 71 (1985); Sun et al., Plants 13 (2024); POWO (2026). = Sumatra and Borneo. = KALIMANTAN. = Lycopodium beccarii Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 6: 29 (1912).
  3. Huperzia bolanica (Rosenst.) Holub, Folia Geobot. Phytotax. 20: 71 (1985). – Phlegmariurus bolanicus (Rosenst.) A.R.Field & Bostock, PhytoKeys 20: 36 (2013); Hassler (2004-onwards). Lycopodium bolanicum Rosenst., Repert. Spec. Nov. Regni Veg. 12: 181 (1913). – Urostachys bolanicus (Rosenst.) Herter ex Nessel, Bärlappgew. 234 (1939). – Huperzia kaysseri (Nessel) Holub, Folia Geobot. Phytotax. 20: 74 (1985). – Urostachys kaysseri Nessel, Bärlappgew. 31, t. 4, f. 10 (1939) (nom. inval.), et Rev. Südam. Bot. 6: 156 (1940). Endemic to New Guinea. PAPUA.
  4. Huperzia campestris (Alderw.) Holub, Folia Geobot. Phytotax. 20: 71 (1985); Sun et al., Plants 13 (2024); POWO (2026). = New Guinea. = PAPUA. = Lycopodium campestre Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 3 (1917).
  5. Huperzia carinata (Desv. ex Poir.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 247 (1875). – Phlegmariurus carinatus (Poir.) Ching, Acta Bot. Yunnan. 4: 120 (1982); Hassler (2004-onwards). – Huperzia albescens (F.M.Bailey) Holub, Folia Geobot. Phytotax. 20: 70 (1985). – Huperzia laxa (C.Presl) T.Sen & U.Sen, Fern Gaz. 11(6): 420, f. 3, a-j (1978) (non (Desv.) Khokr.). – Huperzia laxa (C.Presl) Trevis., Atti Soc. Ital. Sci. Nat. 17: 247 (1875). – Huperzia struthioloides (C.Presl) Rothm., Fedde, Repert. Spec. Nov. 54: 61 (1944). – Huperzia vanikorensis (Copel.) Holub, Folia Geobot. Phytotax. 20: 77 (1985). – Lycopodium acrostachyum Hook. & Grev., Icon. Fil. 2: t. 181 (1831). – Lycopodium albescens F.M.Bailey, Queensl. Dept. Agric. Brisbane Bot. Bull. 14: 16 (1896). – Lycopodium carinatum Desv. ex Poir., Encycl. suppl. 3: 555 (1813 (1814)). – Lycopodium carinatum var. minus Tagawa, Acta Phytotax. Geobot. 15: 7 (1964). – Lycopodium laxum C.Presl, Rel. Haenk. 1(1): 83 (1825). – Lycopodium pendulum Roxb., Calcutta J. Nat. Hist. 4: 472 (1844). – Lycopodium struthioloides C.Presl, Rel. Haenk. 1: 82 (1825). – Lycopodium urostachyum Wall. ex Ridl., J. Straits Branch Roy. Asiat. Soc. 80: 142 (1919). – Lycopodium vanikorense Copel., J. Arnold Arbor. 12: 49 (1931). – Phlegmariurus laxus (C.Presl) Satou, Hikobia 12(3): 268 (1997). – Selaginella struthioloides (C.Presl) Underw., Bull. Torrey Bot. Cl. 25: 132 (1898). – Urostachys albescens (F.M.Bailey) Herter, Rev. Südam. Bot. 8: 86 (1949). – Urostachys carinatus (Poir.) Herter, Bot. Arch. 3: 15 (1923). – Urostachys struthioloides (C.Presl) Herter ex Nessel, Bärlappgew. 170 (1939). – Urostachys vanikorensis (Copel.) Herter, Index Lyc. 87 (1949). China, Taiwan, Nicobar, Malesia, Australia, to Pacific. SUMATRA, KALIMANTAN, SULAWESI, PAPUA.
  6. Huperzia ceylanica (Spring) Rothm., Fedde, Repert. Spec. Nov. 54: 59 (1951). – Phlegmariurus ceylanicus (Spring) comb. ined, Hassler (2004-onwards). – Huperzia vorwerkii (Nessel) Holub, Folia Geobot. Phytotax. 20: 78 (1985). – Lycopodium ceylanicum Spring, Mém. Acad. Roy. Sci. Belg. 15 (1): 37 (1843), 24(2): 16 (1850). – Urostachys ceylanicus (Spring) Herter ex Nessel, Bärlappgew. 52 (1939). – Urostachys vorwerkii Nessel, Fedde, Repert. Spec. Nov. 36(12-15): 178 (1934). Sri Lanka, India, China, Sumatra. SUMATRA.
  7. Huperzia copelandiana (Chou & Bartlett) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Phlegmariurus copelandianus (R.C.Y.Chou & Bartlett) A.R.Field, PhytoKeys 109: 64 (2018); Hassler (2004-onwards). – Lycopodium petiolatum Copel., Univ. Calif. Publ. Bot. 14: 377 (1929) (non Herter (1909)). – Lycopodium copelandianum R.C.Y.Chou & Bartlett, Bull. Torrey Bot. Cl. 74: 369 (1947). Sumatra. SUMATRA.
  8. Huperzia copelandii (Herter) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Phlegmariurus filicaulon (Copel.) A.R.Field & Testo, Molec. Phylogen. Evol. 94(B): 645 (2015); Hassler (2004-onwards). – Lycopodium filicaulon Copel., Phil. Journ. Sci. 60: 100, t. 2 (1936) (non Lycopodium filicaule Hook. fil. (1844)). – Urostachys copelandii Herter, Est. Bot. Reg. Uruguaya (Index Lycop.) 20: 57 (1949). New Guinea and Solomon Islands. PAPUA.
  9. Huperzia coralium (Spring) Holub, Folia Geobot. Phytotax. 20: 71 (1985). – Phlegmariurus coralium (Spring) A.R.Field & Bostock, PhytoKeys 20: 37 (2013); Hassler (2004-onwards). – Lycopodium coralium Spring, Miq., Pl. Jungh. 3: 273 (1854). – Urostachys coralium (Spring) Herter ex Nessel, Bärlappgew. 234 (1939). Peninsular Malaysia (Perak); Java, Solomon Isl. JAVA.
  10. Huperzia crebra (Alderw.) Holub, Folia Geobot. Phytotax. 20: 71 (1985). – Phlegmariurus creber (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 37 (2013); Hassler (2004-onwards). – Lycopodium crebrum Alderw., Bull. Jard. Bot. Buitenz. II, 24: 4 (1917 (crebre)). – Urostachys creber (Alderw.) Herter ex Nessel, Bärlappgew. 181 (1939). New Guinea. PAPUA.
  11. Huperzia dalhousieana (Spring) Trevis., Atti Soc. Ital. Sci. Nat. 17: 247 (1875). – Phlegmariurus dalhousieanus (Spring) A.R.Field & Bostock, PhytoKeys 20: 38 (2013); Hassler (2004-onwards). – Urostachys dalhousianus (Spring) Herter ex Nessel, Bärlappgew. 185 (1939). – Lycopodium dalhousieanum Spring, Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 25 (1849)). – Huperzia caudifolia (Alderw.) Holub, Folia Geobot. Phytotax. 20: 71 (1985). – Urostachys caudifolius (Alderw.) Herter ex Nessel, Bärlappgew. 185 (1939). – Lycopodium caudifolium Alderw., Bull. Jard. Bot. Buitenz. II, 1: 14 (1911). – Huperzia clarae (F.M.Bailey) Holub, Folia Geobot. Phytotax. 20: 71 (1985). – Lycopodium clarae F.M.Bailey, Queensl. Dept. Agric. Brisbane Bot. Bull. 21 (Bot. Bull. 7): 69 (1893). – Urostachys clarae (F.M.Bailey) Herter ex Nessel, Bärlappgew. 185 (1939). – Lycopodium glaucum Ces., Atti Acad. Napoli 7: 35 (1876). – Urostachys glaucus (Ces.) Herter, Rev. Südam. Bot. 8: 86 (1949). – Lycopodium ingens A.Gepp ex Nessel, Bärlappgew. 185 (1939), nom. nud. – Huperzia magnifica (Brownlie) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Lycopodium magnificum Brownlie, Beih. Nova Hedwigia 55: 22-23, t. 2e, f. 3-4 (1977). Peninsular Malaysia, Borneo, Maluku, Australia, Pacific. MALUKU.
  12. Huperzia dielsii (Herter) Holub, Folia Geobot. Phytotax. 20: 72 (1985). – Phlegmariurus dielsii (Herter) A.R.Field & Bostock, PhytoKeys 20: 39 (2013); Hassler (2004-onwards). – Lycopodium dielsii Herter, Bot. Jahrb. Syst. 54: 229 (1916). – Urostachys dielsii (Herter) Herter ex Nessel, Bärlappgew. 213 (1939). Endemic to New Guinea. PAPUA.
  13. Huperzia everettii (Herter) Holub, Folia Geobot. Phytotax. 20: 72 (1985); Sun et al., Plants 13 (2024); POWO (2026). = Endemic to Sulawesi: South Sulawesi. = Lycopodium everettii Herter in Beibl. Bot. Jahrb. Syst. 98: 32, 43 (1909).
  14. Huperzia filiformis (Sw.) Holub, Folia Geobot. Phytotax. 20: 72 (1985). – Phlegmariurus filiformis (Sw.) W.H.Wagner, Contr. Univ. Michigan Herb. 20: 242 (1995); Hassler (2004-onwards). – Huperzia polytrichoides (Kaulf.) Trevis., Bärlappgew. 129 (1939). – Huperzia verticillata (L.fil.) Trevis., Atti Soc. Critt. Ital. 17: 248 (1875). – Huperzia verticillata var. filiformis (Sw.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1875). – Lycopodium acerosum var. filiforme (Sw.) Spring, Mart., Fl. Bras. 1(2): 111 (1840). – Lycopodium filiforme Sw., Schrad., J. Bot. 1800(2): 114 (1801) (Syn. Fil. 174, 398, t. 4, f. 3 (1806)). – Lycopodium polytrichoides Kaulf., Enum. Fil. 6 (1824). – Lycopodium portoricense Underw. & F.E.Lloyd, Bull. Torrey Bot. Cl. 33: 108 (1906). – Lycopodium setaceum Lam., Encycl. 3: 653 (1789 (1792)). – Lycopodium squarrulosum Desv., (1823). – Lycopodium tortile Christ, Bull. Soc. bot. Genève 1(5): 235 (1909). – Lycopodium verticillatum L.fil., Suppl. Pl. 448 (1782). – Lycopodium verticillatum var. filiforme (Sw.) Spring, Mém. Acad. Roy. Sci. Bruxelles 12: 111 (1840). – Phlegmariurus verticillatus (L.fil.) A.R.Field & Testo, Molec. Phylogen. Evol. 94(B): 645 (2015). – Plananthus setaceus (Lam.) P.Beauv., Prodr. aethéogam. 112 (1805). – Plananthus verticillatus (L.fil.) P.Beauv., Prodr. aethéogam. 112 (1805). – Stachygynandrum verticillatum (L.fil.) P.Beauv., Prodr. aethéogam. 113 (1805). – Urostachys filiformis (Sw.) Herter, Est. Bot. Reg. Uruguaya (Index Lycop.) 20: 61 (1949). – Urostachys polytrichoides (Kaulf.) Herter ex Nessel, Bärlappgew. 129 (1939). – Urostachys portoricensis (Underw. & F.E.Lloyd) Herter, Repert. Spec. Nov. Regni Veg. 19(11-13): 163 (1923). – Urostachys verticillatus (L.fil.) Herter, Beih. Bot. Centralbl. 39: 249 (1922). – Urostachys verticillatus var. filiformis (Sw.) Herter ex Nessel & Hoehne, Arch. Bot. Sao Paulo 1: 404 (1927). – Urostachys verticillatus var. gracilis Herter ex Nessel, Bärlappgewächse 123 (1939). Old World. JAVA, KALIMANTAN, SULAWESI, MALUKU, PAPUA.
  15. Huperzia flagellacea (Kuhn) Holub, Folia Geobot. Phytotax. 20: 72 (1985). – Phlegmariurus flagellaceus (Kuhn) A.R.Field & Bostock, PhytoKeys 20: 40 (2013); Hassler (2004-onwards). – Lycopodium flagellaceum Kuhn, Forschungsr. Gazelle 4: 15 (1889). – Urostachys flagellaceus (Kuhn) Herter ex Nessel, Bärlappgew. 237 (1939). Endemic to New Guinea. PAPUA.
  16. Huperzia gedeana (Alderw.) Holub, Folia Geobot. Phytotax. 20: 73 (1985); Sun et al., Plants 13 (2024); POWO (2026). = Endemic to Java. = JAVA. = Lycopodium gedeanum Alderw. in Malayan Ferns Fern Allies, Suppl. 1(Corr.): 65 (1917). – Urostachys gedeanus (Alderw.) Herter ex Nessel in Bärlappgewächse: 45 (1939).
  17. Huperzia goebelii (Nessel) Holub, Folia Geobot. Phytotax. 20: 73 (1985). – Phlegmariurus goebelii (Nessel) A.R.Field & Bostock, PhytoKeys 20: 40 (2013); Hassler (2004-onwards). – Huperzia heroldii (Nessel) Holub, Folia Geobot. Phytotax. 20: 73 (1985). – Urostachys goebelii Nessel, Fedde, Repert. 36: 188, t. 175 (1934). – Urostachys heroldii Nessel, Rev. Südam. Bot. 6: 167, t. 14, f. 69 (1940). Sumatra and Borneo. SUMATRA, KALIMANTAN.
  18. Huperzia gunturensis (Alderw.) Holub, Folia Geobot. Phytotax. 20: 73 (1985). – Phlegmariurus gunturensis (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 41 (2013); Hassler (2004-onwards). – Lycopodium gunturense Alderw., Bull. Jard. Bot. Buitenz. II, 1: 14 (1911). – Urostachys gunturensis (Alderw.) Herter ex Nessel, Bärlappgew. 183 (1939). Endemic to Java. JAVA.
  19. Huperzia hellwigii (Warb.) Holub, Folia Geobot. Phytotax. 20: 73 (1985). – Phlegmariurus hellwigii (Warb.) A.R.Field & Bostock, PhytoKeys 20: 41 (2013); Hassler (2004-onwards). – Lycopodium hellwigii Warb., Monsunia 1: 97 (1898). – Urostachys hellwigii (Warb.) Herter ex Nessel, Bärlappgew. 230 (1939). Endemic to New Guinea. PAPUA.
  20. Huperzia hippuris (Desv. ex Poir.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1875). – Phlegmariurus hippuris (Desv. ex Poir.) A.R.Field & Testo, Molec. Phylogen. Evol. 94(B): 645 (2015); Hassler (2004-onwards). – Lycopodium blumeanum de Vriese, Tijdschr. Nat. Gesch. 3: 338 (1844). – Lycopodium hippuris Desv. ex Poir., Encycl. suppl. 3: 559 (1813 (1814)). – Urostachys hippuris (Poir.) Herter, Index Lyc. 65 (1949). Java, Philippines?, Maluku, Samoa?. JAVA.
  21. Huperzia horizontalis (Nessel) Holub, Folia Geobot. Phytotax. 20: 73 (1985). – Phlegmariurus horizontalis (Nessel) A.R.Field & Bostock, PhytoKeys 20: 41 (2013); Hassler (2004-onwards). – Lycopodium aellenii (Herter) Tagawa, Acta Phytotax. Geobot. 26: 108 (1974). – Lycopodium horizontale Alderw., Bull. Jard. Bot. Buitenz. II, 7: 30 (1912) (non C. Presl (1825)). – Urostachys horizontalis Herter ex Nessel, Bärlappgew. 225 (1939). Endemic to Borneo. KALIMANTAN.
  22. Huperzia javanica (Sw.) C.Y.Yang, Bull. Acad. Military Med. Sci. 13: 368 (1989); Sun et al., Plants 13 (2024); POWO (2026). = Tropical Asia and West Pacific. = SUMATRA, JAVA, KALIMANTAN, SULAWESI, NUSA TENGGARA, MALUKU. = Huperzia longipetiolata (Spring) Y.C.Yang in Chin. Tradit. Herbal Drugs 16(7): 32 (1982), nom. superfl. – Huperzia serrata subsp. javanica (Sw.) Satou in Hikobia 12: 268 (1997). – Huperzia serrata f. longipetiolata (Spring) Ching in Acta Bot. Yunnan. 3: 294 (1981). – Huperzia serrata var. longipetiolata (Spring) H.M.Chang in Taiwania 46: 378 (2001). – Lycopodium javanicum Sw. in J. Bot. (Schrader) 1800(2): 114 (1801). – Lycopodium serratum var. javanicum (Sw.) Makino in Bot. Mag. (Tokyo) 12: 13 (1898), nom. superfl. – Lycopodium serratum var. longipetiolatum Spring in Mém. Acad. Roy. Sci. Belgique 24: 18 (1849). – Phlegmariurus javanicus (Sw.) H.F.Wang, X.Y.Dong & K.P.Ma in Checkl. Vasc. Pl. E. Asia 1: 4 (2025), without basionym ref. – Urostachys javanicus (Sw.) Herter in Estud. Bot. Reg. Uruguay 20: 66 (1949).
  23. Huperzia keysseri (Herter & Schltr. ex Diels) P.J.Edwards, Guide Alpine Subalpine Fl. Mount Jaya (R. J. Johns & al.) 31 (2006). – Phlegmariurus keysseri (Herter & Schltr. ex Diels) comb.ined, Hassler (2004-onwards). – Lycopodium keysseri Herter & Schltr. ex Diels, Bot. Jahrb. Syst. 62: 456 (1929). Endemic to New Guinea (Irian Jaya, Papua New Guinea). PAPUA.
  24. Huperzia lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) Holub, Folia Geobot. Phytotax. 20: 74 (1985). – Phlegmariurus lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) A.R.Field & Bostock, PhytoKeys 20: 42 (2013); Hassler (2004-onwards). – Lycopodium lauterbachii E.Pritz. ex K.Schum. & Lauterb., K. Schum. & Lauterb., Fl. Schutzgeb. Südsee 149 (1900). – Urostachys lauterbachii (K.Schum. & Lauterb.) Herter ex Nessel, Bärlappgew. 203 (1939). Endemic to New Guinea. PAPUA.
  25. Huperzia ledermannii (Herter) Holub, Folia Geobot. Phytotax. 20: 74 (1985). – Phlegmariurus ledermannii (Herter) A.R.Field & Bostock, PhytoKeys 20: 42 (2013); Hassler (2004-onwards). – Lycopodium ledermannii Herter, Bot. Jahrb. Syst. 54: 232 (1916). – Lycopodium phlegmaria var. brachystachyum Rosenst., Repert. Spec. Nov. Regni Veg. 12: 530 (1913). – Urostachys ledermannii (Herter) Herter ex Nessel, Bärlappgew. 236 (1939). Sulawesi and New Guinea. PAPUA.
  26. Huperzia macgregorii (Baker) Holub, Folia Geobot. Phytotax. 20: 74 (1985). – Phlegmariurus macgregorii (Baker) A.R.Field & Bostock, PhytoKeys 20: 42 (2013); Hassler (2004-onwards). – Huperzia versteegii (Alderw.) Holub, Folia Geobot. Phytotax. 20: 77 (1985). – Huperzia whartoniensis (Nessel) Holub, Folia Geobot. Phytotax. 20: 78 (1985). – Lycopodium macgregorii Baker, J. Bot. 28: 109 (1890). – Lycopodium versteegii Alderw., Bull. Jard. Bot. Buitenz. II, 24: 4 (1917). – Urostachys macgregorii (Baker) Herter ex Nessel, Bärlappgew. 184 (1939). – Urostachys versteegii (Alderw.) Herter ex Nessel, Bärlappgew. 182 (1939). – Urostachys whartoniensis Nessel, Fedde, Repert. 39: 62, t. 190b (1935). New Guinea and Solomon Islands. PAPUA.
  27. Huperzia miniata (Spring) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1874 publ. 1875); Sun et al., Plants 13 (2024); POWO (2026). = Java, Philippines, and Sulawesi. = JAVA, SULAWESI. = Lycopodium miniatum Spring in Bull. Acad. Roy. Sci. Bruxelles 8(2): 513 (1841). – Lycopodium selago var. miniatum (Spring) B.C.Tan & Tolentino in Philipp. J. Sci. 116: 437 (1987).
  28. Huperzia minutifolia (Alderw.) Holub, Folia Geobot. Phytotax. 20: 74 (1985). – Phlegmariurus minutifolius (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 43 (2013); Hassler (2004-onwards). – Lycopodium coralium var. minutifolium Alderw., Mal. Fern-Allies 44 (1915). – Urostachys minutifolius (Alderw.) Herter, Index Lyc. 70 (1949). – Lycopodium minutifolium (Alderw.) Alderw., Bull. Jard. Bot. Buitenz. II, 28: 44 (1918). Endemic to New Guinea. PAPUA.
  29. Huperzia multifaria (Alderw.) Holub, Folia Geobot. Phytotax. 20: 75 (1985). – Phlegmariurus multifarius (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 43 (2013); Hassler (2004-onwards). – Lycopodium multifarium Alderw., Bull. Jard. Bot. Buitenz. III, 5: 226 (1922). – Urostachys multifarius (Alderw.) Herter ex Nessel, Bärlappgew. 207 (1939). Endemic to Maluku. MALUKU.
  30. Huperzia nummularifolia (Blume) Jermy, Brit. Fern Gaz. 10: 176 (1971). – Phlegmariurus nummularifolius (Blume) Ching, Acta Bot. Yunnan. 3: 298 (1981); Hassler (2004-onwards). – Lycopodium nummularifolium Blume, Enum. Pl. Jav. 2: 263 (1828). – Urostachys nummularifolius (Blume) Herter, Phil. Journ. Sci.22: 182 (1923). – Lycopodium rotundifolium Roxb., Calcutta J. Nat. Hist. 4: 473 (1844). Malesia to Vanuatu. SUMATRA, JAVA, KALIMANTAN, SULAWESI, MALUKU.
  31. Huperzia patentissima (Alderw.) Holub, Folia Geobot. Phytotax. 20: 75 (1985). – Phlegmariurus patentissimus (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 45 (2013); Hassler (2004-onwards). – Lycopodium patentissimum Alderw., Bull. Jard. Bot. Buitenz. II, 24: 4 (1917). – Urostachys patentissimus (Alderw.) Herter ex Nessel, Bärlappgew. 226 (1939). Seram and New Guinea. MALUKU, PAPUA.
  32. Huperzia phlegmaria (L.) Rothm., Fedde, Repert. Spec. Nov. 54: 62 (1944). – Phlegmariurus phlegmaria (L.) Holub, Preslia 36(1): 21 (1964); Hassler (2004-onwards). – Huperzia feejeensis (Luerss.) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Huperzia phlegmaria var. divaricatum Blume, Enum. Pl. Jav. 262 (1828). – Huperzia simonii (Nessel) Holub, Folia Geobot. Phytotax. 20: 76 (1985). – Lepidotis phlegmaria (L.) P.Beauv., Prodr. aethéogam. 109 (1805). – Lycopodium filiforme Roxb., Calcutta J. Nat. Hist. 4: 473 (1844) (non Sw. (1800)). – Lycopodium horizontale var. submoniliforme Alderw., Bull. Jard. Bot. Buitenz. II, 24: 4 (1917). – Lycopodium phlegmaria L., Sp. Pl. 2: 1101 (1753). – Lycopodium phlegmaria var. longibracteatum Domin, Bibl. Bot. (1928). – Lycopodium phlegmaria var. marchionicum E.D.Br., (1931). – Lycopodium phlegmarium var. feejeense Luerss., Fil. Graeff. 1: 270 (1871). – Lycopodium transiens Alderw., Bull. Jard. Bot. Buitenz. III, 5: 227 (1922). – Lycopodium vrieseanum Spring, Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 33 (1849). – Phlegmariurus phlegmaria (L.) T.Sen & U.Sen, Fern Gaz. 11 (6): 421, f. 4 a (1978) (isonym). – Urostachys borneensis Herter, Phil. Journ. Sci. 22: 181 (1923). – Urostachys feejeensis (Luerss.) Herter, Notul. Syst. (Paris) 15: 355 (1958). – Urostachys leonis Herter, Index Lyc. 67 (1949), nom. nov. for U. borneensis Herter – Urostachys phlegmaria (L.) Herter ex Nessel, Bärlappgew. 215 (1939). – Urostachys simonii Nessel, Rev. Südam. Bot. 6: 167, excl. fig. (1940). – Urostachys submoniliformis (Alderw) Herter, Index Lyc. 84 (1949). – Urostachys transiens (Alderw.) Herter ex Nessel, Bärlappgew. 143 (1939). – Urostachys vrieseanus (Spring) Herter ex Nessel, Bärlappgew. 236 (1939). Old World tropics. SUMATRA, JAVA, KALIMANTAN, SULAWESI, NUSA TENGGARA, MALUKU, PAPUA.
  33. Huperzia phlegmarioides (Gaudich.) Rothm., Fedde, Repert. Spec. Nov. 54: 62 (1944). – Phlegmariurus phlegmarioides (Gaudich.) A.R.Field & Bostock, PhytoKeys 20: 45 (2013); Hassler (2004-onwards). – Huperzia kandavuensis (Nessel) Holub, Folia Geobot. Phytotax. 20: 74 (1985). – Huperzia pseudophlegmaria (Kuhn) Holub, Folia Geobot. Phytotax. 20: 76 (1985). – Lycopodium phlegmarioides Gaudich., Voy. Uranie [Freycinet] 1: 281, t. 23 (1828 (16 Aug)). – Lycopodium pseudophlegmaria Kuhn, Forschungsr. Gazelle 4: 16 (1889). – Lycopodium tetrapterygium (Herter) F.M.Bailey, Proc. Roy. Soc. Queensl. 1: 150 (1884). – Urostachys kandavuensis Nessel, Fedde, Repert. 39: 67, t. 192a (1935). – Urostachys phlegmarioides (Gaudich.) Herter ex Nessel, Bärlappgew. 247 (1939). – Urostachys pseudophlegmaria (Kuhn) Herter ex Nessel, Bärlappgew. 237 (1939). – Urostachys tetrapterygius (F.M.Bailey) Herter, Index Lyc. 86 (1949). – Phlegmariurus pseudophlegmaria (Kuhn) A.R.Field & Testo, Molec. Phylogen. Evol. 94(B): 645 (2015). Peninsular Malaysia, Maluku, New Guinea, Australia, and Pacific. MALUKU, PAPUA.
  34. Huperzia phyllantha (Hook. & Arn.) Holub, Folia Geobot. Phytotax. 20: 75 (1985). – Phlegmariurus phyllanthus (Hook. & Arn.) R.D.Dixit, J. Bombay Nat. Hist. Soc. 77 (3): 541 (1980 [1981]); Hassler (2004-onwards). – Huperzia macrostachys (Spring) Holub, Folia Geobot. Phytotax. 20: 74 (1985). – Huperzia phyllantha (Hook. & Arn.) Panigrahi, Higher Plants Indian Subcont., add. ser. Indian J. Forest. 7, 4: 109 (1993), isonym. – Lycopodium macrostachys Spring, Mém. Acad. Roy. Sci. Belg. 24 (Monogr. Lyc. 2): 30 (1849). – Lycopodium pachystachyon Spring, Bull. Acad. Roy. Soc. Bruxelles 8: 518 (1841) (non Desv. ex Poir. (1814)). – Lycopodium pachystachyum var. phyllanthum (Hook. & Arn.) Hillebr., Fl. Hawaii Isl. 644 (1888). – Lycopodium phyllanthum Hook. & Arn., Bot. Beech. Voy. 102 (1832 (Oct-Nov.)). – Phlegmariurus macrostachys (Hook. ex Spring) N.C.Nair & S.R.Ghosh, J. Econ. Taxon. Bot. 12(1): 194 (1988). – Urostachys macrostachys (Spring) Herter ex Nessel, Bärlappgew. 211 (1939). – Urostachys pachystachyon (Spring) Herter ex Nessel, Bärlappgew. 211 (1939). – Urostachys pachystachyon (Spring) Rothm., Fedde, Repert. Spec. Nov. 54: 62 (1944). – Urostachys phyllanthum (Hook. & Arn.) Herter, Phil. Journ. Sci. 22: 181 (1923). S Asia to Malesia and Pacific. KALIMANTAN, PAPUA.
  35. Huperzia pinifolia (Blume) Trevis., Atti Soc. Ital. Sci. Nat. 17: 247 (1874). – Phlegmariurus pinifolius (Trevis.) Kiew, PhytoKeys 96: 108 (2018); Hassler (2004-onwards). – Huperzia piscium (Herter) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Lycopodium clavatum var. hochreutineri Nessel, Bärlappgew. 300, t, f. 46 (1939) (nom. inval.). – Lycopodium pinifolium Blume, Enum. Pl. Jav. 2: 264 (1828) (non Kaulf. (1824)). – Lycopodium pinifolium var. hochreutineri Herter, Ann. Jard. Cons. Bot. Genève 15/16: 227 (1912). – Lycopodium piscium (Herter) Tagawa & K.Iwats., Acta Phytotax. Geobot. 22: 103 (1967). – Urostachys hochreutineri (Herter) Herter ex Nessel, Bärlappgew. 234 (1939). – Urostachys mercurii Herter, Index Lyc. 69 (1949). – Urostachys pinifolius (Blume) Herter, Bot. Arch. 3: 16 (1923). – Urostachys piscium Herter, Index Lyc. 75 (1949). Thailand, Malesia, to Pacific. SUMATRA, JAVA, KALIMANTAN, SULAWESI, MALUKU.
  36. Huperzia prolifera (Blume) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1875). – Phlegmariurus proliferus (Blume) A.R.Field & Bostock, PhytoKeys 20: 46 (2013); Hassler (2004-onwards). – Lycopodium proliferum Blume, Enum. Pl. Jav. 2: 265 (1828). – Urostachys prolifer (Blume) Herter, Index Lyc. 77 (1949). Java and Kalimantan east to Solomon Islands. JAVA, KALIMANTAN, SULAWESI, NUSA TENGGARA, MALUKU.
  37. Huperzia pulcherrima (Wall. ex Hook. & Grev.) Pic.Serm., Webbia 24: 719 (1970). – Phlegmariurus pulcherrimus (Hook. & Grev.) Á. & D.Löve, Taxon 26: 324 (1977); Hassler (2004-onwards). – Huperzia pulcherrima (Wall. ex Hook. & Grev.) T.Sen & U.Sen, Fern Gaz. 11 (6): 419 (1978), isonym – Huperzia setacea Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1875). – Huperzia setacea (Trevis.) Rothm., Feddes Repert. Spc. Nov. Regn. Veg. 54: 61 (1961), isonym – Huperzia subulifolia (Wall. ex Hook. & Grev.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1875). – Huperzia subulifolia var. assamica J.P.Mandal & U.Sen, Biol. Mem. 3(2): 225 (1979). – Huperzia taiwanensis (Ching) Holub, Folia Geobot. Phytotax. 20(1): 77 (1985). – Lycopodium pulcherrimum Wall. ex Hook. & Grev., Icon. Fil. 1(2): t. 38 (1827). – Lycopodium setaceum Buch.-Ham. ex D.Don, Prodr. Fl. Nepal. 18 (1825) (nom. illeg., non Lycopodium setaceum Lam. (1789)). – Lycopodium setaceum var. pulcherrimum (Wall. ex Hook. & Grev.) C.B.Clarke, Trans. Linn. Soc. Lond. 2, Bot. 1: 590 (1880). – Lycopodium subulifolium Wall. ex Hook. & Grev., Icon. Fil. t. 49 (1831) (fide Fraser-Jenkins 2008). – Phlegmariurus subulifolius (Wall. ex Hook. & Grev.) S.R.Ghosh, Pterid. Fl. E. India 66 (2004), et J. Econ. Taxon. Bot. 33(2): 364 (2009). – Phlegmariurus taiwanensis Ching, Acta Bot. Yunnan. 4: 124 (1982 (nom. nov. superfl. for Lycopodium pulcherrimum Hook. & Grev.). – Urostachys amphorae Herter, Index Lyc. 51 (1949 (nom. nov. superfl. for Urostachys pulcherrimus (Hook. & Grev.) Nessel)). – Urostachys pulcherrimus (Wall. ex Hook. & Grev.) Herter ex Nessel, Bärlappgew. 72 (1939). – Urostachys setaceus (Buch.-Ham. ex D.Don) Herter ex Nessel, Arch. Bot. Sao Paulo 1: 408 (1927) (Bärlappgewächse 141 (1939)). – Urostachys subulifolius (Wall. ex Hook. & Grev.) Herter, Index Lyc. 84 (1949). Mainland S and SE Asia to Borneo and Sulawesi. KALIMANTAN, SULAWESI.
  38. Huperzia rupicola (Alderw.) Holub, Folia Geobot. Phytotax. 20: 76 (1985); Sun et al., Plants 13 (2024); POWO (2026). = New Guinea. = PAPUA. = Lycopodium rupicola Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 39 (1914). – Phlegmariurus rupicola (Alderw.) A.R.Field & Bostock in PhytoKeys 20: 46 (2013).
  39. Huperzia salvinioides (Herter) Holub, Folia Geobot. Phytotax. 20. 76 (1985). – Phlegmariurus salvinioides (Herter) Ching, Acta Bot. Yunnan. 4: 122 (1982); Hassler (2004-onwards). – Lycopodium salvinioides (Herter) Tagawa, Acta Phytotax. Geobot. 8: 20 (1939). – Urostachys formosanus Herter ex Nessel, Bärlappgew. 229 (1939) (nom. inval.), Rev. Südam. Bot. 6: 167, excl. fig. (1940). – Urostachys salvinioides Herter, Bot. Arch. 3: 18 (1923). – Lycopodium edanoi Copel., Phil. Journ. Sci. 46: 209 (1931). – Huperzia edanoi (Copel.) Holub, Folia Geobot. Phytotax. 20: 72 (1985). – Urostachys edanoi (Copel.) Nessel, Bärlappgew. 230 (1939). East Asia, Borneo, Philippines. KALIMANTAN.
  40. Huperzia setifolia (Alderw.) Holub, Folia Geobot. Phytotax. 20: 76 (1985). – Phlegmariurus setifolius (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 47 (2013); Hassler (2004-onwards). – Lycopodium setifolium Alderw., Bull. Jard. Bot. Buitenz. II, 16: 40 (1914). – Urostachys setifolius (Alderw.) Herter ex Nessel, Bärlappgew. 233 (1939). Endemic to Borneo. KALIMANTAN.
  41. Huperzia squarrosa (G.Forst.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 247 (1875). – Phlegmariurus squarrosus (G.Forst.) Á. & D.Löve, Taxon 26: 324 (1977); Hassler (2004-onwards). – Huperzia acutifolia (Desv ex Poir.) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Huperzia epiceifolia (Desv. ex Poir.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1875). – Huperzia forsteri Poir., nom. illeg. superfl. – Huperzia hookeri (Hook. & Grev.) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Huperzia remoganense Hayata – Huperzia squarrosa (G.Forst.) Rothm., Fedde, Repert. Spec. Nov. 54: 62 (1944), comb. superfl. – Huperzia ulicifolia (Sw.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1875). – Lycopodium acutifolium Desv. ex Poir., Encycl. suppl. 3: 559 (1813 publ. 1814). – Lycopodium epiceifolium Desv. ex Poir., Encycl. suppl. 3: 559 (1813 (1814)). – Lycopodium forsteri Poir., Encycl. suppl. 3: 545 (1813 (1814)) (nom. superfl.). – Lycopodium gnidioides Blanco, Fl. Filip. 824 (1837), ed. 2.569 (1845), ed. 3 (3): 239 (1879). – Lycopodium hookeri Wall. ex Hook. & Grev., Icon. Fil. 2: t. 185 (1831). – Lycopodium lohitense D.D.Pant & P.S.Pandey, (as ´lohitensis´, fide Fraser-Jenkins et al. 2015). – Lycopodium madagascariense Desv., (nom. inval.). – Lycopodium pseudosquarrosum Pamp., Bull. R. Soc. Tosc. Orticult. III, 13: 99, t. 2 (1908). – Lycopodium squarrosum G.Forst., Fl. ins. austr. Prodr. 479 (1786). – Lycopodium ulicifolium Sw., Syn. Fil. 177 (1806). – Phlegmariurus acutifolius (Desv. ex Poir.) A.R.Field & Testo, Molec. Phylogen. Evol. 94(B): 644 (2015). – Phlegmariurus ulicifolius (Sw.) S.R.Ghosh, Pterid. Fl. E. India 64 (2004). – Plananthus squarrosa (Sw.) P.Beauv., Prodr. aethéogam. 112 (1805). – Urostachys epiceifolius (Desv. ex Poir.) Herter ex Nessel, Bärlappgew. 202 (1939). – Urostachys epiceifolius var. acutifolius (Desv.) Herter ex Nessel, Bärlappgew. 292 (1939). – Urostachys madagascariensis (Desv. ex Nessel) Herter, Est. Bot. Reg. Uruguaya (Index Lycop.) 20: 68 (1949), Alt. title: Index Lycopodiarum 68 (1949). – Urostachys madagascariensis (Nessel) Herter, Index Lyc. 68 (1949). – Urostachys squarrosus (G.Forst.) Herter, Bot. Arch. 3: 14 (1923). – Urostachys squarrosus f. madagascariensis Desv. ex Nessel, Revista Sudamericana de Botánica 6,5/6: 165, 166, t. 12, fig. 60 (1940). – Urostachys squarrosus var. proliferus (Blume) Herter ex Nessel, Bärlappgew. 200 (1939). – Urostachys ulicifolius (Sw.) Herter ex Nessel, Bärlappgew. 204 (1939). S and SE Asia, Malesia, Australia, and Pacific. SUMATRA, JAVA, KALIMANTAN, SULAWESI, NUSA TENGGARA, MALUKU.
  42. Huperzia subfalciformis (Alderw.) Holub, Folia Geobot. Phytotax. 20: 77 (1985). – Phlegmariurus subfalciformis (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 47 (2013); Hassler (2004-onwards). – Lycopodium subfalciforme Alderw., Bull. Jard. Bot. Buitenz. II, 28: 44 (1918). – Urostachys subfalciformis (Alderw.) Herter ex Nessel, Bärlappgew. 227 (1939). – Phlegmariurus subfalciformis (Alderw.) A.R.Field & Testo, Molec. Phylogen. Evol. 94(B): 645 (2015), comb. superfl. Endemic to New Guinea. PAPUA.
  43. Huperzia sumatrana (Alderw.) Holub, Folia Geobot. Phytotax. 20: 77 (1985); Sun et al., Plants 13 (2024); POWO (2026). = Endemic to Sumatra. = SUMATRA. = Lycopodium sumatranum Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 44 (1918). – Urostachys sumatranus (Alderw.) Herter ex Nessel in Bärlappgewächse: 66 (1939).
  44. Huperzia talamauana (Alderw.) Holub, Folia Geobot. Phytotax. 20: 77 (1985). – Phlegmariurus talamauanus (Alderw.) A.R.Field & Bostock, PhytoKeys 20: 48 (2013); Hassler (2004-onwards). – Lycopodium talamauanum Alderw., Bull. Jard. Bot. Buitenz. II,28: 45 (1918). – Urostachys talamauanus (Alderw.) Herter ex Nessel, Bärlappgew. 212 (1939). Endemic to Sumatra. SUMATRA.
  45. Huperzia tauri (Herter) Holub, Folia Geobot. Phytotax. 26(1): 92 (1991). – Phlegmariurus tauri (Herter) A.R.Field & Testo, Molec. Phylogen. Evol. 94(B): 645 (2015); Hassler (2004-onwards). – Lycopodium apiculatum Spring, Miq., Pl. Jungh. 3: 272 (1854) (non Poir. 1814). – Urostachys tauri Herter, Index Lyc. 85 (1949). – Urostachys apiculatus (Spring) Herter ex Nessel, Bärlappgew. 227 (1939). – Lycopodium tauri (Herter) M.Kato, Acta Phytotax. Geobot. 39(4-6): 137 (1988). Java, Sulawesi, Maluku. JAVA, SULAWESI, MALUKU.
  46. Huperzia terrae-guilelmii (Herter) Holub, Folia Geobot. Phytotax. 20: 77 (1985). – Phlegmariurus terrae-guilelmii (Herter) A.R.Field & Bostock, PhytoKeys 20: 48 (2013); Hassler (2004-onwards). – Lycopodium terrae-guilelmii Herter, Bot. Jahrb. Syst. 54: 229 (1916). – Urostachys terrae-guilelmii (Herter) Herter ex Nessel, Bärlappgew. 231 (1939). Endemic to Indonesian New Guinea. PAPUA.
  47. Huperzia tetrasticha (Kunze) Holub, Folia Geobot. Phytotax. 20: 77 (1985). – Phlegmariurus tetrastichus (Kunze) A.R.Field & Bostock, PhytoKeys 20: 48 (2013); Hassler (2004-onwards). – Lycopodium tetrastichum Kunze, Bot. Zeit. (Berlin) 6: 99 (1848). – Lycopodium sarasinorum Christ, Verh. Naturf. Ges. Basel 11: 34, t. 2o, f. 11-12 (1894). – Urostachys sarasinorum (Christ) Herter ex Nessel, Bärlappgew. 182 (1939). – Urostachys tetrastichus (Kunze) Herter ex Nessel, Bärlappgew. 183 (1939, by reference to Alderw. (1915a: 35)). Peninsular Malaysia, Java, Kalimantan, Sulawesi, Maluku. KALIMANTAN, SULAWESI, MALUKU.
  48. Huperzia verticillata (L.f.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248 (1874 publ. 1875); Sun et al., Plants 13 (2024); POWO (2026). = Tropics and subtropics regions of the world. = KALIMANTAN. = Lycopodium verticillatum L.f. in Suppl. Pl.: 448 (1782). – Phlegmariurus verticillatus (L.f.) A.R.Field & Testo in Molec. Phylogen. Evol. 94(B): 645 (2015). – Plananthus verticillata (L.f.) P.Beauv. in Prodr. Aethéogam.: 112 (1805). – Urostachys verticillatus (L.f.) Herter in Beih. Bot. Centralbl., Abt. 2. 39: 249 (1922). – Stachygynandrum verticillatum (L.f.) P.Beauv. in Prodr. Aethéogam.: 113 (1805). – Huperzia filiformis (Sw.) Holub in Folia Geobot. Phytotax. 20: 72 (1985). – Huperzia polytrichoides (Kaulf.) Trevis. in Atti Soc. Ital. Sci. Nat. 17: 248 (1874 publ. 1875). – Lycopodium acerosum var. filiforme (Sw.) Spring in C.F.P.von Martius & auct. suc. (eds.), Fl. Bras. 1(2): 111 (1840). – Lycopodium filiforme Sw. in J. Bot. (Schrader) 1800(2): 114 (1801). – Lycopodium polytrichoides Kaulf. in Enum. Filic.: 6 (1824). – Lycopodium portoricense Underw. & F.E.Lloyd in Bull. Torrey Bot. Club 33: 108 (1906). – Lycopodium setaceum Lam. in Encycl. 3: 653 (1792). – Lycopodium tortile Christ in Bull. Soc. Bot. Genève, sér. 2, 1: 235 (1909). – Lycopodium verticillatum var. apertum Fée in Crypt. Vasc. Brésil 1: 222 (1869). – Lycopodium verticillatum var. filiforme (Sw.) Spring in Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 47 (1842). – Phlegmariurus filiformis (Sw.) W.H.Wagner in Contr. Univ. Michigan Herb. 20: 242 (1995). – Urostachys portoricensis Herter in Repert. Spec. Nov. Regni Veg. 19: 163 (1923). – Urostachys verticillatus var. filiformis Herter ex Nessel & Hoehne in Arch. Bot. São Paulo 1: 399 (1927). – Urostachys verticillatus var. gracilis Herter ex Nessel in Revista Sudamer. Bot. 6: 163 (1940).
  49. Huperzia zollingeri (Herter) Holub, Folia Geobot. Phytotax. 20: 78 (1985); Sun et al., Plants 13 (2024); POWO (2026). = Sumatra and Java. = SUMATRA, JAVA. = Lycopodium zollingeri Herter in Beibl. Bot. Jahrb. Syst. 98: 34, 48 (1909). – Huperzia monticola (Alderw.) Holub in Folia Geobot. Phytotax. 20: 75 (1985). – Lycopodium monticola Alderw. in Malayan Ferns Fern Allies, Suppl. 1(Corr.): 65 (1917).

 

LYCOPODIELLA Holub

  1. Lycopodiella caroliniana (L.) Pic.Serm., Webbia 23: 165 (1968); Sun et al., Plants 13 (2024); POWO (2026). = Pantropical. = PAPUA. = Lepidotis caroliniana (L.) P.Beauv. in Prodr. Aethéogam.: 108 (1805). – Lycopodium carolinianum L. in Sp. Pl.: 1104 (1753). – Pseudolycopodiella caroliniana (L.) Holub in Folia Geobot. Phytotax. 18: 442 (1983). = ≡ var. caroliniana. = As the species. = PAPUA.
  2. Lycopodiella cernua (L.) Pic.Serm., Webbia 23: 166 (1968); Sun et al., Plants 13 (2024); POWO (2026). = Tropics and subtropics regions of the world. = SUMATRA, JAVA, KALIMANTAN, SULAWESI, NUSA TENGGARA, MALUKU, PAPUA. = Lepidotis cernua (L.) P.Beauv. in Prodr. Aethéogam.: 108 (1805). – Lycopodium cernuum L. in Sp. Pl.: 1103 (1753). – Palhinhaea cernua (L.) Franco & Vasc. in Bol. Soc. Brot., sér. 2, 41: 25 (1967). – Lycopodiella cernua var. caboverdeana (Gilli) O.Eriksson, A.Hansen & Sunding in Fl. Macaronesia, ed. 2, 1: 92 (1979). – Lycopodiella cernua var. capillacea (Willd. ex Spring) D.Heller in Consp. Fl. Orient. 9: 167 (1994). – Lycopodiella cernua var. sikkimensis (Müll.Hal.) Panigrahi & Sarn.Singh in Ferns Fern-Allies Arunachal Pradesh 1: 54 (2005). – Lycopodium brevibracteatum Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 5 (1917). – Lycopodium capillaceum (Willd. ex Spring) Willd. ex Spring in Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 79 (1842). – Lycopodium cernuum var. blancheanum Stehlé in Fl. Agron. Antilles Franç. 3: 138 (1960). – Lycopodium cernuum var. brevifolium Christ in Bull. Herb. Boissier, sér. 2, 3: 620 (1903), nom. nud. – Lycopodium cernuum var. caboverdeanum Gilli in Verh. Zool.-Bot. Ges. Wien 115: 39 (1976). – Lycopodium cernuum var. capillaceum Willd. ex Spring in Flora 21: 165 (1838). – Lycopodium cernuum var. chaminadeanum Stehlé in Fl. Agron. Antilles Franç. 3: 138 (1960). – Lycopodium cernuum var. crassifolium Spring in Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 80 (1842). – Lycopodium cernuum f. durissimum Herter ex Nessel in Bärlappgewächse: 355 (1939). – Lycopodium cernuum var. dussii Christ in Bot. Jahrb. Syst. 24: 147 (1897). – Lycopodium cernuum f. ingens B.C.Tan & Tolentino in Philipp. J. Sci. 116: 438 (1987). – Lycopodium cernuum var. laxum Blume in Enum. Pl. Javae 2: 266 (1828). – Lycopodium cernuum f. macrostachyum Christ in Primit. Fl. Costaric. 3: 55 (1901). – Lycopodium cernuum var. majus C.Presl in Reliq. Haenk. 1: 80 (1825). – Lycopodium cernuum var. medium C.Presl in Reliq. Haenk. 1: 81 (1825). – Lycopodium cernuum var. minus C.Presl in Reliq. Haenk. 1: 81 (1825). – Lycopodium cernuum var. panamense Wercklé ex Nessel in Bärlappgewächse: 366 (1939). – Lycopodium cernuum f. sikkimense (Müll.Hal.) H.S.Kung in Acta Phytotax. Sin. 18: 239 (1980). – Lycopodium cernuum var. sikkimense (Müll.Hal.) C.B.Clarke in Trans. Linn. Soc. London, Bot. 1: 592 (1880). – Lycopodium cernuum var. watsonianum Herter ex Nessel in Bärlappgewächse: 366 (1939). – Lycopodium heeschii Müll.Hal. in Bot. Zeitung (Berlin) 19: 164 (1861). – Lycopodium hupeanum Müll.Hal. in Bot. Zeitung (Berlin) 19: 165 (1861). – Lycopodium marianum Willd. in Sp. Pl., ed. 4. 5: 31 (1810). – Lycopodium polycephalum Colenso in Trans. & Proc. New Zealand Inst. 27: 401 (1894 publ. 1895). – Lycopodium secundum Müll.Hal. in Bot. Zeitung (Berlin) 19: 164 (1861). – Lycopodium sikkimense Müll.Hal. in Bot. Zeitung (Berlin) 19: 164 (1861). – Palhinhaea brevibracteata (Alderw.) Holub in Folia Geobot. Phytotax. 20: 78 (1985). – Palhinhaea capillacea (Willd. ex Spring) Holub in Folia Geobot. Phytotax. 20: 78 (1985). – Palhinhaea cernua f. ciliatomarginata W.M.Chu in Fl. Yunnanica 20: 718 (2006). – Palhinhaea cernua var. lanceolata Gias & Pasha in Indian Fern J. 14: 26 (1998), without latin descr. or type. – Palhinhaea cernua f. laxa (Blume) Satou in Hikobia 12: 269 (1997). – Palhinhaea cernua f. sikkimensis (Müll.Hal.) H.S.Kung in Fl. Sichuanica 6: 50 (1988). – Palhinhaea cernua var. sikkimensis (Müll.Hal.) Ching in Acta Bot. Yunnan. 3: 6 (1981). – Palhinhaea crassifolia (Spring) Fraser-Jenk. & Kholia in Annot. Checkl. Ind. Pterid. 3: 310 (2020). – Palhinhaea lufengensis C.Y.Yang in Tradit. Herbal Drugs 16(7): 34 (1985). – Palhinhaea mariana (Willd.) Holub in Folia Geobot. Phytotax. 20: 79 (1985). – Palhinhaea polycephala (Colenso) Holub in Folia Geobot. Phytotax. 20: 79 (1985).
  3. Lycopodiella hydrophila (Alderw.) P.J.Edwards in R.J.Johns & al., Guide Alpine Subalpine Fl. Mount Jaya: 35 (2006); Sun et al., Plants 13 (2024); POWO (2026). = New Guinea. = PAPUA. = Lycopodium hydrophilum Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 4 (1917).
  4. Lycopodiella tomentosa (Alderw.) P.J.Edwards in R.J.Johns & al., Guide Alpine Subalpine Fl. Mount Jaya: 35 (2006); Sun et al., Plants 13 (2024); POWO (2026). = New Guinea. = PAPUA. = Lycopodium tomentosum Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 6 (1917). – Palhinhaea tomentosa (Alderw.) Holub in Folia Geobot. Phytotax. 20: 79 (1985).

 

LYCOPODIUM L.

  1. Lycopodium angustiramosum (Alderw.) Herter, Estud. Bot. Reg. Uruguay 20: 4 (1949); Sun et al., Plants 13 (2024); POWO (2026). = New Guinea and Solomon Islands. = PAPUA. = Diphasiastrum angustiramosum (Alderw.) Holub in Preslia 47: 107 (1975). – Lycopodium complanatum angustiramosum Alderw. in Bull. Jard. Bot. Buitenzorg, ser. 2, 24: 6 (1917).
  2. Lycopodium casuarinoides Spring, Bull. Acad. Roy. Sci. Bruxelles 8: 521 (1841); Sun et al., Plants 13 (2024); POWO (2026). = South to East Asia and Malesia. = SUMATRA, KALIMANTAN, PAPUA. = Diphasium casuarinoides (Spring) J.P.Mandal & U.Sen in Biol. Mem. 3: 252 (1979), without basionym reef. – Lepidotis casuarinoides (Spring) Rothm. in Feddes Repert. Spec. Nov. Regni Veg. 54: 67 (1944). – Lycopodiastrum casuarinoides (Spring) Holub ex R.D.Dixit in J. Bombay Nat. Hist. Soc. 77: 541 (1980 publ. 1981). – Lycopodium comansf. in Fl. Antarct. 1: 112 (1844). – Lycopodium filicaule Hook.f. in Fl. Antarct. 1: 112 (1844). – Lycopodium jacobsonii Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 23: 22 (1916). – Lycopodium penicilliferum Alderw. in Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 40 (1914).
  3. Lycopodium clavatum, Sp. Pl.: 1101 (1753); Sun et al., Plants 13 (2024); POWO (2026). = Almost throughout the World. = KALIMANTAN, SULAWESI, MALUKU, PAPUA. = Lepidotis clavata (L.) P.Beauv. in Prodr. Aethéogam.: 108 (1805). = ≡ subsp. clavatum = Almost throughout the world. = KALIMANTAN, SULAWESI, MALUKU, PAPUA. = Lepidotis ciliata P.Beauv. in Prodr. Aethéogam.: 108 (1805). – Lepidotis inflexa P.Beauv. in Prodr. Aethéogam.: 109 (1805). – Lycopodium aristatum Humb. & Bonpl. ex Willd. in Sp. Pl., ed. 4. 5: 17 (1810). – Lycopodium aristatum var. desvauxianum (Spring) Spring in C.F.P.von Martius & auct. suc. (eds.), Fl. Bras. 1(2): 114 (1840). – Lycopodium aristatum var. incurvum Hook. & Grev. in Bot. Misc. 2: 376 (1831). – Lycopodium aristatum var. robustius Grev. & Hook. in Bot. Misc. 2: 376 (1831). – Lycopodium ciliatum (P.Beauv.) Sw. in Syn. Fil. (Kiliae): 179 (1806), nom. illeg. homonym. post. – Lycopodium clavatum var. aristatum (Humb. & Bonpl. ex Willd.) Spring in Flora 21: 173 (1838). – Lycopodium clavatum var. asiaticum Ching in Acta Bot. Yunnan. 4: 224 (1982). – Lycopodium clavatum var. borbonicum Bory in Voy. îles Afrique 2: 205 (1804). – Lycopodium clavatum f. brevipedunculatum Louis-Marie in Rev. Oka Agron. Inst. Agric. 14: 144 (1940). – Lycopodium clavatum f. decapitatum Louis-Marie in Rev. Oka Agron. Inst. Agric. 14: 144 (1940). – Lycopodium clavatum var. desvauxianum Spring in Flora 21: 173 (1838). – Lycopodium clavatum var. distachyum Spring in Flora 21: 171 (1838). – Lycopodium clavatum var. equisetoides Schwacke in Arch. Bot. São Paulo 1: 438 (1927). – Lycopodium clavatum var. eriostachys (Fée) Nessel & Hoehne in Arch. Bot. São Paulo 1: 435 (1927). – Lycopodium clavatum f. frondescens Vict. in Contr. Lab. Bot. Univ. Montréal 3: 25 (1925). – Lycopodium clavatum f. frondosum Vict. in Contr. Lab. Bot. Univ. Montréal 3: 25 (1925). – Lycopodium clavatum f. incurvum (Hook. & Grev.) Nessel in Bärlappgewächse: 305 (1939). – Lycopodium clavatum var. inflexum (P.Beauv.) Spring in Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 90 (1842). – Lycopodium clavatum var. jamaicense Spring in Flora 21: 174 (1838). – Lycopodium clavatum var. laurentianum Vict. in Contr. Lab. Bot. Univ. Montréal 3: 23 (1925). – Lycopodium clavatum var. minarum Christ in C.A.W.Schwacke, Pl. Nov. Mineir. 2: 41 (1900). – Lycopodium clavatum var. montanum Stehlé in Mém. Soc. Bot. France 1953-1954: 44 (1954). – Lycopodium clavatum var. nataliense Nessel in Repert. Spec. Nov. Regni Veg. 36: 191 (1934). – Lycopodium clavatum var. paucidivisum Spring in Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 91 (1842). – Lycopodium clavatum var. piliferum (Raddi) Nessel & Hoehne in Arch. Bot. São Paulo 1: 435 (1927), nom. superfl. – Lycopodium clavatum var. preslianum Spring in Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 90 (1842). – Lycopodium clavatum var. raddianum Spring in Flora 21: 172 (1838). – Lycopodium clavatum f. robustinus Nessel & Hoehne in Arch. Bot. São Paulo 1: 439 (1927). – Lycopodium clavatum var. robustius (Grev. & Hook.) Nakai in Bot. Mag. (Tokyo) 39: 197 (1925). – Lycopodium clavatum var. serpens (C.Presl) Spring in Flora 21: 172 (1838). – Lycopodium clavatum var. sitchaense Spring in Flora 21: 173 (1838). – Lycopodium clavatum f. sterilis House in Bull. New York State Mus. Nat. Hist. 243-244: 19 (1923). – Lycopodium clavatum var. subremotum Vict. in Contr. Lab. Bot. Univ. Montréal 3: 24 (1925). – Lycopodium clavatum var. trichiatum (Bory) Nessel in Bärlappgewächse: 31 (1939). – Lycopodium clavatum var. trichophyllum (Desv.) Nessel & Hoehne in Arch. Bot. São Paulo 1: 436 (1927), nom. superfl. – Lycopodium clavatum f. trilobum (Farw.) M.Broun in Index No. Amer. Ferns: 107 (1938). – Lycopodium clavatum subvar. trilobum Farw. in Amer. Fern J. 27: 18 (1937). – Lycopodium eriostachys Fée in Crypt. Vasc. Brésil 1: 224 (1869). – Lycopodium inflexum (P.Beauv.) Sw. in Syn. Fil. (Kiliae): 179 (1806). – Lycopodium officinale Neck. in Meth. Mus.: 150 (1771). – Lycopodium piliferum Raddi in Pl. Bras. Nov. Gen. 1: 79 (1825). – Lycopodium preslii Hook. & Grev. in Bot. Misc. 2: 377 (1831). – Lycopodium serpens C.Presl in Reliq. Haenk. 1: 81 (1825), nom. illeg. homonym. post. – Lycopodium torridum Gaudich. in Voy. Uranie: 283 (1828). – Lycopodium trichiatum Bory in Voy. îles Afrique 1: 350 (1804). – Lycopodium trichiatum var. desvauxianum (Spring) Spring in Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 92 (1842). – Lycopodium trichophyes Spreng. in Syst. Veg., ed. 16. 4(1): 13 (1827). – Lycopodium trichophyllum Desv. in Mém. Soc. Linn. Paris 6: 184 (1827). – Urostachys plutonis Herter in Estud. Bot. Reg. Uruguay 20: 75 (1949).
  4. Lycopodium hygrophilum, Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 5 (1917); Sun et al., Plants 13 (2024); POWO (2026). = New Guinea. = PAPUA.
  5. Lycopodium japonicum, Syst. Veg. ed. 14. 944: (1784); Hassler (2004-onwards). – Lycopodium centrochinense Ching, Acta Bot. Yunnan. 4(3): 224-225 (1982) (fide Fraser-Jenkins 2008, 2017). – Lycopodium clavatum var. nipponicum Nakai, Bot. Mag. 39(463): 197 (1925). – Lycopodium divaricatum Wall. ex Hook. & Grev., Bot. Misc. 2: 377 (1831). – Lycopodium clavatum var. wallichianum Spring, Monogr. Lyc. 1: 90 (1842). – Lycopodium interjectum Ching & H.S.Kung, Acta Bot. Yunnan. 4(3): 221 (1982). – Lycopodium japonicum f. yunnanense (Ching) W.M.Chu, Fl. Yunnan. 20: 25 (2006). – Lycopodium kinabaluense Ching, Acta Bot. Yunnan. 4(3): 223 (1982). – Lycopodium pseudoclavatum Ching, Acta Bot. Yunnan. 4(3): 222-223 (1982) (fide Fraser-Jenkins 1997). – Lycopodium pseudoclavatum var. yunnanense Ching, Acta Bot. Yunnan. 4(3): 223 (1982) (fide Fraser-Jenkins 1997). – Lycopodium simulans Ching & H.S.Kung, Acta Bot. Yunnan. 4(3): 220-221 (1982). – Lycopodium taliense Ching, Acta Bot. Yunnan. 4: 221 (1982). – Stachygynandrum japonicum (Thunb.) P.Beauv., Prodr. aethéogam. 113 (1805). S Asia, China, mainland SE Asia, Malesia, to Solomon Islands. KALIMANTAN, SULAWESI, MALUKU, PAPUA.
  6. Lycopodium scariosumForst., Fl. Ins. Austr.: 87 (1786); Sun et al., Plants 13 (2024); POWO (2026). = Borneo, Wallacea, and southern Australia. = KALIMANTAN, SULAWESI. = Diphasium scariosum (G.Forst.) Rothm. in Feddes Repert. Spec. Nov. Regni Veg. 54: 65 (1944). – Diphasium decurrens (R.Br.) Holub in Folia Geobot. Phytotax. 20: 78 (1985). – Lycopodium decurrens R.Br. in Prodr. Fl. Nov. Holland.: 165 (1810). – Lycopodium scariosum var. decurrens (R.Br.) Hook. in Cent. Ferns: t. LXVI (1854).
  7. Lycopodium volubileForst., Fl. Ins. Austr.: 86 (1786); Sun et al., Plants 13 (2024); POWO (2026). = Malesia. = SUMATRA, JAVA, KALIMANTAN, SULAWESI, NUSA TENGGARA, MALUKU, PAPUA. = Pseudodiphasium volubile (G.Forst.) Holub in Folia Geobot. Phytotax. 18: 442 (1983).
  8. Lycopodium wightianum ex Hook. & Grev., Bot. Misc. 2: 379 (1831); Sun et al., Plants 13 (2024); POWO (2026). = South Asia and Malesia. = KALIMANTAN, PAPUA. = Diphasiastrum wightianum (Wall. ex Hook. & Grev.) Holub in Preslia 47: 108 (1975). – Diphasium wightianum (Wall. ex Hook. & Grev.) Pic.Serm. in Webbia 24: 720 (1970). – Stachygynandrum wightianum (Wall. ex Hook. & Grev.) C.Presl in Abh. Königl. Böhm. Ges. Wiss., ser. 5, 3: 583 (1845). – Diphasiastrum novoguineense (Nessel) Holub in Preslia 47: 108 (1975). – Lycopodium alpinum var. novoguineense Nessel in Repert. Spec. Nov. Regni Veg. 39: 69 (1936). – Lycopodium novoguineense (Nessel) Herter in Estud. Bot. Reg. Uruguay 20: 30 (1949).